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価格はGLS400d 4MATICが1263万円、上級グレードのGLS580 4MATIC スポーツが1669万円。GLS580は4リッターV8ツインターボにISG（インテグレーテッド・スターター・ジェネレーター）を組み合わせたマイルドハイブリッド仕様となる。
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前後のトルク配分を0〜100%まで連続可変させることで、オン/オフの走りを両立させる新開発の4WD（4MATIC）を搭載する。オフロード用のローレンジギアを備えたオフロード エンジニアリング パッケージをオプションで用意した。
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インパネには12.3インチの大型ディスプレイが2画面並び、クラシック/スポーツ/プログレッシブ/ジェントルから表示モードが選択できる。
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AIRMATICサスペンションには、コーナリング時にボディを内側に傾けることで乗員が感じる遠心力低減させるダイナミックカーブ機能（CURVE）が採用された。
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ラゲッジのサイドには2/3列目シートを電動で折り畳めるスイッチが備わっており、ラゲッジ側からのシートアレンジが楽におこなえる。ラゲッジ容量は355〜2400リッター。
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